the.com/tusk

a tooth so ambitious it left the mouth and started running the whole face

means a long, pointed tooth that protrudes well beyond the mouth, as in elephants, walruses, and boars.

from From Old English 'tusc' or 'tux,' related to Old Frisian 'tusk' meaning much the same thing. Beyond that the trail goes cold — its deeper roots are uncertain, possibly tangled with words for 'tooth,' which would make sense for a tooth that grew too big for its job.

true teeth
Tusks are overgrown incisors or canines, not horns
never stop
Elephant tusks grow throughout their entire lives
handedness
Elephants favor one tusk like we favor a hand
narwhal twist
That spiral tusk is a sensor-packed tooth piercing the lip
walrus winch
Walruses haul their bodies onto ice using tusks
for instance

elephant ivory trade — approximately 20,000 elephants killed annually for tusks, primarily in africa

walrus hunting alaska — indigenous iñupiat and yupik peoples harvest walrus tusks for cultural and subsistence purposes

mammoth tusk fossil trade — siberian permafrost yields thousands of tusks yearly, sold in international markets since 1990s

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